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Calvin Cycle (n.)
1.(MeSH)The synthesis by organisms of organic chemical compounds, especially carbohydrates, from carbon dioxide using energy obtained from light rather than from the oxidation of chemical compounds. Photosynthesis comprises two separate processes: the light reactions and the dark reactions. In higher plants; GREEN ALGAE; and CYANOBACTERIA; NADPH and ATP formed by the light reactions drive the dark reactions which result in the fixation of carbon dioxide. (from Oxford Dictionary of Biochemistry and Molecular Biology, 2001)
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Calvin Cycle (n.) (MeSH)
Calvin-Benson-Bassham Cycle (MeSH), Calvin-Benson Cycle (MeSH), Carbon Fixation, Photosynthetic (MeSH), Dark Reactions of Photosynthesis (MeSH), Photosynthesis (MeSH), Reductive Pentose Phosphate Cycle (MeSH)
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Calvin Cycle (n.) [MeSH]
Physiology, Plant, Plant Physiology - Metabolic Networks, Metabolic Networks and Pathways, Metabolic Pathways - Phototrophic Growth, Phototrophic Process, Phototrophic Processes, Phototrophy - Carbohydrate Metabolism, Metabolism, Carbohydrate - Photochemistry[Hyper.]
Chlorophyll, Chlorophyll 740 - Chloroplast, Chloroplasts, Etioplasts[Analogie]
Wikipedia
The Calvin cycle, Calvin–Benson-Bassham (CBB) cycle, reductive pentose phosphate cycle or C3 cycle is a series of biochemical redox reactions that take place in the stroma of chloroplasts in photosynthetic organisms. It is also known as the dark reactions.
The cycle was discovered by Melvin Calvin, James Bassham, and Andrew Benson at the University of California, Berkeley[1] by using the radioactive isotope carbon-14. It is one of the light-independent (dark) reactions used for carbon fixation.
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Photosynthesis occurs in two stages. In the first stage, light-dependent reactions capture the energy of light and use it to make the energy-storage molecules ATP and NADPH. The light-independent Calvin cycle uses the energy from short-lived electronically excited carriers to convert carbon dioxide and water into organic compounds[2] that can be used by the organism (and by animals that feed on it). This set of reactions is also called carbon fixation. The key enzyme of the cycle is called RuBisCO. In the following biochemical equations, the chemical species (phosphates and carboxylic acids) exist in equilibria among their various ionized states as governed by the pH.
The enzymes in the Calvin cycle are functionally equivalent to many enzymes used in other metabolic pathways such as gluconeogenesis and the pentose phosphate pathway, but they are to be found in the chloroplast stroma instead of the cell cytoplasm, separating the reactions. They are activated in the light (which is why the name "dark reaction" is misleading), and also by products of the light-dependent reaction. These regulatory functions prevent the Calvin cycle from being respired to carbon dioxide. Energy (in the form of ATP) would be wasted in carrying out these reactions that have no net productivity.
The sum of reactions in the Calvin cycle is the following:
Hexose (six-carbon) sugars are not a product of the Calvin cycle. Although many texts list a product of photosynthesis as C6H12O6, this is mainly a convenience to counter the equation of respiration, where six-carbon sugars are oxidized in mitochondria. The carbohydrate products of the Calvin cycle are three-carbon sugar phosphate molecules, or "triose phosphates," namely, glyceraldehyde-3-phosphate (G3P).
In the first stage of the Calvin cycle, a CO2 molecule is incorporated into one of two three-carbon molecules (glyceraldehyde 3-phosphate or G3P), using up two molecules of ATP and two molecules of NADPH, both of which were produced in the light-dependent stage. Three steps are involved.
The next stage in the Calvin cycle is to regenerate RuBP. Five G3P molecules produce three RuBP molecules, using up three molecules of ATP. Since each CO2 molecule produces two G3P molecules, three CO2 molecules produce six G3P molecules, of which five are used to regenerate RuBP, leaving a net gain of one G3P molecule per three CO2 molecules (as would be expected from the number of carbon atoms involved).
The regeneration stage can be broken down into steps.
Thus, of six G3P produced, five are used to make three RuBP (5C) molecules (totaling 15 carbons), with only one G3P available for subsequent conversion to hexose. This requires nine ATP molecules and six NADPH molecules per three CO2 molecules. The equation of the overall Calvin cycle is shown diagrammatically below.
RuBisCO also reacts competitively with O2 instead of CO2 in photorespiration. The rate of photorespiration is higher at high temperatures. Photorespiration turns RuBP into 3-PGA and 2-phosphoglycolate, a 2-carbon molecule that can be converted via glycolate and glyoxalate to glycine. Via the glycine cleavage system and tetrahydrofolate, two glycines are converted into serine +CO2. Serine can be converted back to 3-phosphoglycerate. Thus, only 3 of 4 carbons from two phosphoglycolates can be converted back to 3-PGA. It can be seen that photorespiration has very negative consequences for the plant, because, rather than fixing CO2, this process leads to loss of CO2. C4 carbon fixation evolved to circumvent photorespiration, but can occur only in certain plants native to very warm or tropical climates, for example, corn.
The immediate products of one turn of the Calvin cycle are 2 glyceraldehyde-3-phosphate (G3P) molecules, 3 ADP, and 2 NADP+. (ADP and NADP+ are not really "products." They are regenerated and later used again in the Light-dependent reactions). Each G3P molecule is composed of 3 carbons. In order for the Calvin cycle to continue, RuBP (ribulose 1,5-bisphosphate) must be regenerated. So, 5 out of 6 carbons from the 2 G3P molecules are used for this purpose. Therefore, there is only 1 net carbon produced to play with for each turn. To create 1 surplus G3P requires 3 carbons, and therefore 3 turns of the Calvin cycle. To make one glucose molecule (which can be created from 2 G3P molecules) would require 6 turns of the Calvin cycle. Surplus G3P can also be used to form other carbohydrates such as starch, sucrose, and cellulose, depending on what the plant needs.[5]
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